Nighttime levels are adjusted in the Wii version to allow greater use of the Wii Remote, such that players may control the arms of the Werehog by using the Wii Remote and Nunchuk in order to grab onto objects and proceed in the levels. Collecting rings in these levels replenishes the Werehog's health, while a special move, Unleashed Mode, allows the Werehog to channel his energy into increasing the power of his attacks for a short time. Gameplay here sees the player use a variety of melee and combo attacks to defeat large numbers of enemies, collect items, and move items around to advance through the level, and use his stretchable arms to cling to distant objects simple puzzle elements are also featured. Nighttime levels feature slower-paced, action-oriented levels, in which Sonic transforms into the Werehog, a beast form that has great strength and stretchable arms.
Japanese game company Dimps helped design some of the stages. In contrast, the PlayStation 2 and Wii versions of the game represent available Boost using bars, which may be added by performing 'Action Chains', destroying multiple enemies in quick succession through the use of homing attacks, or by collecting rings. The amount of boost remaining may be increased by collecting more rings, and is decreased by using the Boost. In the Xbox 360 and PlayStation 3 versions, an on-screen 'Ring Energy' meter displays how much boost is available.
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The game also features a gameplay mechanic previously used in the Sonic Rush series called the Sonic Boost, which greatly increases Sonic's speed, allowing him to smash through objects, destroy enemies instantly, or even access different level paths. For instance, a new sidestep feature known as the Quick Step is available, allowing Sonic to dodge left and right, and a Drift feature, which allows Sonic to make tighter turns without slowing down.
In addition to moves available in past games, such as the Homing Attack, new moves are also introduced. 2D sections are reminiscent of the Mega Drive/Genesis-era Sonic games, where the player controls Sonic in a side-scrolling fashion, while 3D sections see the camera placed behind Sonic, so the player may move in all directions. Daytime levels focus on Sonic's speed, and to this extent, sees the player control Sonic through fast-moving stages containing both 2D and 3D styles of gameplay.
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